Gyrfalcon

Taxobox
name = Gyrfalcon or Gerfalcon


image_width = 220px
image_caption = Brown morph
status = LC
status_system = iucn3.1
regnum = Animalia
phylum = Chordata
classis = Aves
ordo = Falconiformes
familia = Falconidae
genus = "Falco"
subgenus = ("Hierofalco")
species = "F. rusticolus"
binomial = "Falco rusticolus"
binomial_authority = Linnaeus, 1758
synonyms =
*"Falco candicans"
Gmelin, 1788
*"Falco islandus"
Brünnich, 1764
*"Falco obsoletus"
Gmelin, 1788
*"Falco rusticolus candicans"
Gmelin, 1788
*"Falco rusticolus grebnitzkii"
(Severtzov, 1885)
*"Falco rusticolus intermedius"
Gloger, 1834
*"Falco rusticolus islandus"
Brünnich, 1764 (but see text)
*"Falco rusticolus obsoletus"
Gmelin, 1788
*"Falco rusticolus rusticolus"
Linnaeus, 1758
*"Falco swarthi"
L.H. Miller, 1927
*"Hierofalco grebnitzkii"
Severtzov, 1885)
*"Hierofalco islandus"
(Brünnich, 1764)
*"Hierofalco rusticolus"
(Linnaeus, 1758)
*"Hierofalco rusticolus candicans"
(Gmelin, 1788)

The Gyrfalcon IPAEng|ˈdʒɜrˌfɔlkən or IPAEng|ˈdʒɜrˌfælkən ("Falco rusticolus"), also spelled Gyr Falcon, sometimes Gerfalcon, is the largest of all falcon species. The Gyrfalcon breeds on Arctic coasts and islands of North America, Europe and Asia. It is mainly resident, but some Gyrfalcons disperse more widely after the breeding season, or in winter [Individual vagrancy can take birds for long distances. There is a story in the "Unauthorized Biography of the Spring and Autumn"Verify source|date=August 2007 of a 海东青 (" Hǎidōngqīng": Gyrfalcon) that succumbed to an arrow wound in the garden of Chen Hui GongVerify source|date=August 2007 . Confucius recognized the arrow as one of the Sushen, whose fine stone arrowheads were a famous item of trade and tribute (RAM 2006). Although the Sushen's precise homeland at that time remains unknown, it was in the Manchuria region, no less than c.600 and perhaps more than 1000 km from the Lu capital of Qufu.] .

The bird's common name comes from French "gerfaucon", and in mediaeval Latin is rendered as "gyrofalco". The first part of the word may come from Old High German "gîr" (cf. modern German "Geier"), "vulture", referring to its size compared to other falcons, or the Latin "gȳrus" ("circle", "curved path") from the species' circling as it searches for prey, unlike the other falcons in its range [The Gyrfalcon's names in other Germanic languages provide few clues to resolve this. In Scandinavian languages, it is generally named after its use in falconry, whereas the modern Dutch name "giervalk" is peculiarly ambiguous: "Gier" means "vulture", whereas "gieren" means changing the yaw angle to circle in the air.] . The male gyrfalcon is called a gyrkin in falconry.

Its scientific name is composed of the Latin terms for a falcon, "Falco", and for someone who lives in the countryside, "rusticolus".

Description

This species is a very large falcon, about the same size as the largest buteos. Males are 48 to 61 cm (19 to 24 in) long, weigh 805 to 1350 g (1.8 to 3 lbs) and have a wingspan from 110 to 130 cm (43 to 51 in). Females are rather bulkier and larger at 51 to 65 cm (20 to 26 in) long, a weight of 1180 to 2100 g (2.6 to 4.6 lbs) and have a wingspan ranging from 124 to 160 cm (49 to 64 in). Gyrfalcons are somewhat intermediate between a large Peregrine Falcon and a hawk in general structure; they are unmistakably falcons with pointed wings, but are stockier, broader-winged and longer-tailed than the Peregrine.

Plumage is very variable in this highly polymorphic species: the archetypal morphs are called "white", "silver", "brown" and "black" though coloration spans a continuous spectrum from nearly all-white birds to very dark ones. The brown form of the Gyrfalcon is distinguished from the Peregrine by the cream streaking on the nape and crown and by the absence of a well-defined malar stripe and cap. The black morph has its underside strongly spotted black, not finely barred as in the Peregrine. White form Gyrfalcons are unmistakable, as they are the only predominantly white falcons. Silver birds resemble a light, grey Lanner Falcon of huge size.

There is no difference in coloration between between males and females; juveniles are darker and browner than corresponding adults on average.

ystematics and evolution

The Gyrfalcon is a member of the close-knit hierofalcon complex. In this group, there is ample evidence for rampant hybridization and incomplete lineage sorting which confounds analyses of DNA sequence data to a massive extent; molecular studies with small sample sizes can simply not be expected to yield reliable conclusions in the entire hierofalcon group. The radiation of the entire living diversity of hierofalcons seems to have taken place in the Eemian interglacial at the start of the Late Pleistocene, a mere 130,000-115,000 years ago; the Gyrfalcon seems to represents lineages that expanded into the Holarctic and adapted to local conditions, whereas the inland populations further south, towards northeastern Africa where the radiation probably originated, evolved into the Saker Falcon. Indeed, gyrfalcons hybridize not infrequently with Sakers in the Altay Mountains, and this gene flow seems to be the origin of the "Altai Falcon". [Helbig "et al." (1994), Wink "et al." (1998), Wink "et al." (2004), Nittinger "et al." (2005)]

ubspecies

There is some correlation between locality and the frequency of color morphs. Greenland Gyrfalcons are lightest, with white plumage flecked with grey on the back and wings being most common. Other subpopulations have varying amounts of the darker morphs: the Icelandic birds tend towards pale, and Eurasian ones are considerably darker and not usually have white birds present. Natural separation into regional subspecies is prevented by Gyrfalcons' habit of flying long distances exchanging alleles between subpopulations; thus, the allele distributions for the color polymorphism form clines and in darker birds [The allele combination producing the white morph seems to be recessive.] of unknown origin, theoretically any allele combination might be present. For example, a mating of a pair of captive Gyrfalcons is documented to have produced a clutch of 4 young: one white, one silver, one brown, one black.

In general, geographic variation follows Bergmann's Rule for size and the demands of crypsis for plumage coloration. Several subspecies have been named according to perceived differences between populations ["Falco rusticolus candicans" fron northern Greenland and adjacent North America; "F. r. obsoletus" from the southern Greenland into subarctic North America which is much darker, often brown or black; and "F. r. islandus" (Iceland), "F. r. rusticolus" (Scandinavia inluding the species' type locality, Sweden), as well as "F. r. intermedius" and "F. r. grebnitzkii" (Siberia), which all tend towards more or less dark "silver" coloration (Snow "et al." 1998).] but none of these are consistent and thus no living subspecies are accepted today.

Perhaps the Icelandic population described as "Falco rusticolus islandus" is the most distinct. The predominantly white Arctic forms are parapatric and seamlessly grade into the subarctic populations, whereas the birds of Iceland have presumably less gene flow with their neighbors and indeed show less variation in plumage colors and often look quite similar to a large, washed-out Peregrine Falcon (though their habitus is different). Comprehensive phylogeographic studies are needed to determine the proper status of the Icelandic population however. [White (1994), Snow "et al." (1998)]

There was, however, a paleosubspecies "Falco rusticolus swarthi" during the Late Pleistocene (125,000 - 13,000 years ago). Fossils found in Little Box Elder Cave (Converse County, Wyoming), Dark Canyon Cave (Eddy County, New Mexico) and McKittrick, California were initially described as "Falco swarthi" ("Swarth Falcon" or more properly Swarth's Gyrfalcon) on account of their distinct size. They have meanwhile proven to be largely inseparable from those of living gyrfalcons, except for being somewhat larger. [Miller (1927, 1935), Howard (1971), Emslie (1985)]

Swarth's Gyrfalcon was on the upper end of the present Gyrfalcon's size range, strong females even surpassing it (Miller 1935). It seems to have had some adaptations to the temperate semiarid climate that predominated in its range during the last ice age. Ecologically more similar to the Siberian populations of today (which are generally small birds however) or the Prairie Falcon, this population of temperate steppe habitat must have preyed on landbirds and mammals rather than the water- and seabirds which make up much of American gyrfalcon's diet today.

Ecology

The Gyrfalcon is a bird of tundra and mountains, with cliffs or a few patches of trees. It feeds only on birds and mammals. Like other hierofalcons, it usually hunts in a horizontal pursuit, rather than the Peregrine's speedy stoop from a height. Most prey is killed on the ground, whether they are captured there or, if the victim is a flying bird, forced to the ground. The diet is to some extent opportunistic, but a majority of breeding birds mostly rely on "Lagopus" grouse. Avian prey can range in size from redpolls to geese and can include gulls, corvids, smaller passerines, waders and other raptors (up to the size of "Buteos"). Mammalian prey can range in size from shrews to marmots (sometimes 3 times heavier than the assaulting falcon), and often includes include lemmings, voles, ground squirrels and hares. They only rarely eat carrion.

Reproduction and life history

The Gyrfalcon almost invariably nests on cliff faces. Breeding pairs do not build their own nests, and often use a bare cliff ledge or the abandoned nest of other birds, particularly Golden Eagles and Common Ravens. The clutch can range from 1 to 5 eggs, but is usually 2 to 4. The average size of an egg is 58.46 x 45 mm (2.31 x 1.8 in) and the average weight is 62 g (2.2 oz). The incubation period averages 35 days, until the 52 g (1.8 oz) chicks hatch. The nestlings are brooded usually for 10 to 15 days and leave the nest at 7 to 8 weeks. At 3 to 4 months of age, the immatures become independent of their parents, though they may associate with their siblings through the following winter.

The only natural predator of gyrfalcons are Golden Eagles and even they rarely engage with these formidable falcons. Gyrfalcons have been recorded as aggressively harassing animals that come near their nests, although Common Ravens are the only predators known to successful pick off Gyrfalcon eggs and hatchlings. Even Brown Bears may be dive-bombed, much to their annoyance. Humans, whether accidentally (automobile collisions or poisoning of carrion to kill mammalian scavengers) or intentionally (through hunting), are the leading cause of death for Gyrfalcons. Gyrfalcons that make it to adulthood can live up to 20 years of age.

Relationship with humans

The Gyrfalcon is Iceland's "staðfugl" (national bird), as well as the official bird of Canada's Northwest Territories.

In medieval times, the Gyrfalcon was considered the king's bird. Due to its rarity and the difficulties involved in obtaining it, in falconry the gyrfalcon was generally reserved for kings and nobles. Very seldom was a man of lesser rank seen with a Gyrfalcon on his fist.Fact|date=February 2007

Gyrfalcons are very expensive to buy, and thus owners and breeders may keep them secret to avoid theft. They tend to fly long distances, and falconers may fit a radio-tracker to aid recovery.Fact|date=February 2007

Wild Gyrfalcons are not much exposed to disease, and as a result have weak immune systems. As a result, many gyrfalcons taken from the wild quickly die of disease. Several generations of captive breeding from the survivors causes selection for a stronger immune system and thus better resistance to disease.

Gyrfalcons and the fall of the Liao Dynasty

In the 12th century AD the Jurchen tribes rebelled against the Chinese Liao Dynasty which was set up by Khitan people. The primary cause was that the Khitan nobles extorted a big tax of Gyrfalcons (which are called "Hǎidōngqīng" (海东青) in Chinese), as swan hunting was highly fashionable among the nobility. Especially under the last Liao Emperor Tianzuodi, tax collectors were even entitled to use force to procure the demanded quantity of gyrfalcons. The rebellion caught on, and the Jurchen under chieftain Wanyan Aguda annihilated the Liao empire in 1125, establishing the Jīn Dynasty in its stead. [Beijing Municipal Administration of Cultural Heritage: [http://english.bjww.gov.cn/2006/7-19//18134.shtml Contest for the Southern Capital between the Liao, Song and Jin Dynasties] . Version of 2006-JUL-19. Retrieved 2007-AUG-13.]

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Footnotes

References

*|year=2004|id=49516|title=Falco rusticolus|downloaded=12 May 2006 Database entry includes justification for why this species is of least concern.
* (1985): The late Pleistocene (Rancholabrean) avifauna of Little Box Elder Cave, Wyoming. "Rocky Mountain Geology" 23(2): 63-82. [http://pubs.gg.uwyo.edu/CTGs/CTG_23-2.htm HTML abstract]
* (1994): Phylogenetic relationships among falcon species (genus Falco) according to DNA sequence variation of the cytochrome b gene. "In:" aut|Meyburg, B.-U. & Chancellor, R.D. (eds.): "Raptor conservation today": 593-599. [http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pubwink/1994/4.%201994.pdf PDF fulltext]
* (1971): Quaternary Avian Remains from Dark Canyon Cave, New Mexico. "Condor" 73(2): 237-240. doi|10.2307/1365844 [http://elibrary.unm.edu/sora/Condor/files/issues/v073n02/p0237-p0240.pdf PDF fulltext]
* (1927): The Falcons of the McKittrick Pleistocene. "Condor" 29(3): 150-152. doi|10.2307/1363081 [http://elibrary.unm.edu/sora/Condor/files/issues/v029n03/p0150-p0152.pdf PDF fulltext]
* (1935): A Second Avifauna from the McKittrick Pleistocene. "Condor" 37(2): 72-79. doi|10.2307/1363879 [http://elibrary.unm.edu/sora/Condor/files/issues/v037n02/p0072-p0079.pdf PDF fulltext]
* (2005): Out of Africa? Phylogenetic relationships between "Falco biarmicus" and other hierofalcons (Aves Falconidae). "Journal of Zoological Systematics and Evolutionary Research" 43(4): 321-331. doi|10.1111/j.1439-0469.2005.00326.x [http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pubwink/2005/28.2005.pdf PDF fulltext]
* (2006): " [http://www.royalalbertamuseum.ca/vexhibit/dragon/arrows.htm Dragon Bytes: Did you know about time's arrows?] . Version of October 12, 2006. Retrieved 2007-AUG-13.+
* (1998): "The complete birds of the western Palaearctic on CD-ROM". Oxford University Press. ISBN 0192685791
* (1994): 58. Gyrfalcon. "In:" aut|del Hoyo, Josep; Elliott, Andrew & Sargatal, Jordi (editors): "Handbook of Birds of the World, Volume 2 (New World Vultures to Guineafowl)": 274, plate 28. Lynx Edicions, Barcelona. ISBN 84-87334-15-6
* (1998): Molecular systematics of holarctic raptors (Order Falconiformes). "In:" aut|Chancellor, R.D., Meyburg, B.-U. & Ferrero, J.J. (eds.): "Holarctic Birds of Prey": 29-48. Adenex & WWGBP. [http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pubwink/1998/31.%201998.pdf PDF fulltext]
* (2004): Phylogenetic relationships in the Hierofalco complex (Saker-, Gyr-, Lanner-, Laggar Falcon). "In:" aut|Chancellor, R.D. & Meyburg, B.-U. (eds.): "Raptors Worldwide": 499-504. WWGBP, Berlin. [http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pubwink/2004/27.2004.pdf PDF fulltext]

External links

*http://www.falconryforum.co.uk
*http://www.falconscanada.com/site/Welcome.html
*http://www.kentishfalconry.co.uk/
*http://www.gyrfalcons.co.uk/
*http://www.mosquitonet.com/~akfalconer/ Alaska Falcons
* [http://www.sdakotabirds.com/species/gyrfalcon_info.htm Gyrfalcon Information and Photos] - South Dakota Birds and Birding
* [http://www.birds.cornell.edu/AllAboutBirds/BirdGuide/Gyrfalcon.html Gyrfalcon Species Account] - Cornell Lab of Ornithology
* [http://www.bird-stamps.org/cspecies/3205800.htm Stamps] (with RangeMap)
* [http://vireo.acnatsci.org/search.html?Form=Search&SEARCHBY=Common&KEYWORDS=gyrfalcon&showwhat=images&AGE=All&SEX=All&ACT=All&Search=Search&VIEW=All&ORIENTATION=All&RESULTS=24 Gyrfalcon photo gallery] VIREO