- Climax community
In ecology, a climax community, or climatic climax community, is a biological community of plants and animals which, through the process of ecological succession — the development of vegetation in an area over time — has reached a steady state. This equilibrium occurs because the climax community is composed of species best adapted to average conditions in that area. The term is sometimes also applied in soil development.
The idea of a single climatic climax, which is defined in relation to regional climate, originated with Frederic Clements in the early 1900s. The first analysis of succession as leading to something like a climax was written by Henry Cowles in 1899, but it was Clements who used the term "climax" to describe the idealized endpoint of succession.
Frederic Clements's use of "climax"
Clements described the successional development of an ecological communitiescomparable to the ontogenetic development of individual organisms. Clements suggested only comparisons to very simple organisms. Later ecologists developed this idea that the ecological community is a "superorganism" and even sometimes claimed that communities could be homologous to complex organisms.
Clements's theory sought to define a single climax-type for each area. Arthur Tansley developed this idea with the "polyclimax" -- multiple steady-state end-points, determined by edaphic factors, in a given climatic zone. Clements had called these end-points other tems, not climaxes, and had thought they were not stable, because by definition climax vegetation is best-adapted to the climate of a given area. Henry Gleason's early challenges to Clements's organism simile, and other of his strategies for describing vegetation, were largely disregarded for several decades until substantially vindicated by research in the 1950s and 1960s (below). Meanwhile, climax theory was deeply incorporated in both theoretical ecology and in vegetation management. Clements's terms such as pre-climax, post-climax, plagioclimax and disclimax continued to be used to describe the many communities which persist in states that diverge from the climax ideal for a particular area.
Though the views are sometimes attributed to him, Clements never argued that climax communities must always occur, or that the dominant cause of vegetation is climate, or that the different species in an ecological community are tightly integrated physiologically, or that plant communities have sharp boundaries in time or space. Rather, he employed the idea of a climax community--of the form of vegetation best adapted to some idealized set of environmental conditions--as a conceptual starting point for describing the vegetation in a given area. There are good reasons to believe that the species best adapted to some conditions might appear there, when those conditions occur. But much of Clements's work was devoted to characterizing what happens when those ideal conditions do not occur. In those circumstances, vegetation other than the ideal climax will often occur instead. But those different kinds of vegetation can still be described as deviations from the climax ideal. Therefore Clements developed a very large vocabulary of theoretical terms describing the various possible causes of vegetation, and various non-climax states vegetation adopts as a consequence. His method of dealing with ecological complexity was to define an ideal form of vegetation--the climax community--and describe other forms of vegetation as deviations from that ideal.
Rejection of climax theory
Support among ecologists for the climax theory declined, because they found the theory with its many coined terms difficult to apply, because they were dissatisfied how it compared to observed individual organisms, and because better theories developed.
Although Clements recognized that vegetation follows gradients rather than being tightly bound, his rhetorical comparisons of ecological communities to organisms fostered the impression that communities, including the climax, have distinct edges in space and time. Yet Robert Whittaker's research demonstrated plant species distribute themselves along nutrient and other environmental gradients. Many ecologists saw this as a major reason to stop using the climax concept.
More recent palynological studies showed that modern species assemblages are ephemeral; vegetation in eastern North America since the last glacial maximum has consisted of several different species assemblages, many of which have no analogues in modern "climax" communities. That would mean, at least, that the climax types for those areas could not be stable to the degree Clements believed they were.
Ultimately, even if succession tends towards a steady state, the time required to achieve this state is unrealistically long; in most cases, external disturbances and environmental change occur so frequently that the realization of a climax community is unlikely, and therefore it has come to be regarded as a less useful concept. Long-term vegetation dynamics are now more often characterized as resulting from the action of stochastic factors.
Continuing usage of "climax"
Despite the overall abandonment of climax theory, during the 1990s use of climax concepts again became more popular among some theoretical ecologists. Many authors and nature-enthusiasts continue to use the term "climax" in a diluted form to refer to what might otherwise be called mature or old-growth communities. The term "climax" has also been adopted as description for a late successional stage for marine macroinvertebrate communities.
- ^ Cowles, Henry Chandler. 1899. The Ecological Relations of the Vegetation on the Sand Dunes of Lake Michigan. Botanical Gazette 27(2): 95-117; 27(3): 167-202; 27(4): 281-308; 27(5): 361-391.
- ^ Clements, Frederic E. 1916. Plant Succession: An Analysis of the Development of Vegetation. Washington D.C.: Carnegie Institution of Washington.
- ^ Hagen, Joel B. 1992. An Entangled Bank: The Origins of Ecosystem Ecology. New Brunswick: Rutgers University Press.
- ^ Eliot, Christopher. 2000 . Method and Metaphysics in Clements’s and Gleason’s Ecological Explanations. Studies in History and Philosophy of Biological and Biomedical Sciences 38(1): 85–109.
- ^ Tobey, Ronald C. 1981. Saving the prairies: the life cycle of the founding school of American plant ecology, 1895–1955. Berkeley: University of California Press.
- ^ Whittaker, Robert H. 1953. A consideration of climax theory: the climax as a population and pattern. Ecological Monographs 23: 41–78.
- ^ Cook, James E. 1996. Implications of Modern Successional Theory for Habitat Typing: A Review. Forest Science 42(1): 67–75.
- ^ See, for example, Roughgarden, Jonathan, Robert M. May and Simon A. Levin, editors. 1989. Perspectives in Ecological Theory. Princeton: Princeton University Press.
- ^ Rosenberg R., S. Agrenius, B. Hellman, H. C. Nilsson, and K. Norling. 2002. Recovery of marine benthic habitats and fauna in a Swedish fjord following improved oxygen conditions. Marine Ecology Progress Series 234: 43-53.
Wikimedia Foundation. 2010.
Look at other dictionaries:
climax community — klimaksas statusas T sritis ekologija ir aplinkotyra apibrėžtis Aukščiausia, ilgalaikė ir palyginti pastovi bendrijos (sistemos) raidos pakopa, turinti didelę rūšių įvairovę ir biomasę su sąlygiškai nusistovėjusia produkcija. atitikmenys: angl.… … Ekologijos terminų aiškinamasis žodynas
climax community — The stage in community succession where the community has become relatively stable through successful adjustment to its environment … Geography glossary
climax community — the final, stable, and mature community in a series that appears in succession, which is in equilibrium with the environmental conditions and is composed of a definite group of plant and animal species … Medical dictionary
climax community — /ˌklaɪmæks kəˈmjunəti/ (say .kluymaks kuh myoohnuhtee) noun a stable community of plants composed of the most mesophytic vegetation that the climate can support, and not replaceable by other communities so long as the climate remains unchanged … Australian English dictionary
Plagio climax community — is an area or habitat in which the influences of the human race, have prevented the system from expanding further, an example may be in a beach dune system where the impact of the human race has caused footpath erosion to occur, affecting the… … Wikipedia
Climax — may refer to: Contents 1 Common general uses 2 Brand names and titles 3 Personal name … Wikipedia
Climax species — Climax species, also called late seral, late successional, K selected or equilibrium species, are plant species that will remain essentially unchanged in terms of species composition for as long as a site remains undisturbed. They are the most… … Wikipedia
Climax (narrative) — The death of Caesar in Shakespeare s Julius Caesar is a well known climax The Climax is the point in the story where the main character s point of view changes, or the most exciting/action filled part of the story. It also known has the main… … Wikipedia
community ecology — Introduction study of the organization and functioning of communities (community), which are assemblages of interacting populations of the species living within a particular area or habitat. As populations of species interact with one … Universalium
Climax, Saskatchewan — For other uses, see Climax (disambiguation). Climax, Saskatchewan Village Grain elevators along the railway tracks in Climax … Wikipedia