Semipalmated Sandpiper (Calidris pusilla)
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Aves
Order: Charadriiformes (partim)
  • Charadrii
  • Chionidi
  • Scolopaci
  • Thinocori

Waders, called shorebirds in North America (where "wader" is used to refer to long-legged wading birds such as storks and herons), are members of the order Charadriiformes, excluding the more marine web-footed seabird groups. The latter are the skuas (Stercorariidae), gulls (Laridae), terns (Sternidae), skimmers (Rynchopidae), and auks (Alcidae). Also, the pratincoles (Glareolidae) and the Crab Plover (Dromadidae), which bear greater resemblance to waders, are closely related to the seabirds.

This leaves about 210[verification needed] species, most of which are associated with wetland or coastal environments. Many species of Arctic and temperate regions are strongly migratory, but tropical birds are often resident, or move only in response to rainfall patterns. Some of the Arctic species, such Little Stint are amongst the longest distance migrants, spending the non-breeding season in the southern hemisphere.

The majority of species eat small invertebrates picked out of mud or exposed soil. Different lengths of bills enable different species to feed in the same habitat, particularly on the coast, without direct competition for food. Many waders have sensitive nerve endings at the end of their bills which enable them to detect prey items hidden in mud or soft soil. Some larger species, particularly those adapted to drier habitats will take larger prey including insects and small reptiles.

Many of the smaller species found in coastal habitats, particularly but not exclusively the calidrids, are often named as "sandpipers", but this term does not have a strict meaning, since the Upland Sandpiper is a grassland species.

The smallest member of this group is the Least Sandpiper, small adults of which can weigh as little as 15.5 grams and measure just over 13 cm (5 inches). The largest species is believed to be the Far Eastern Curlew, at about 63 cm (25 inches) and 860 grams (1.9 lb), although the Beach Thick-knee, is the heaviest at about 1 kg (2.2 lb).

In the Sibley-Ahlquist taxonomy, waders and many other groups are subsumed into a greatly enlarged Ciconiiformes order. However, the classification of the Charadriiformes is one of the weakest points of the Sibley-Ahlquist taxonomy, as DNA-DNA hybridization has turned out to be incapable of properly resolving the interrelationships of the group. Formerly, the waders were united in a single suborder Charadrii, but this has turned out to be a "wastebasket taxon", uniting no less than four charadriiform lineages in a paraphyletic assemblage. However, it indicated that the Plains Wanderer actually belonged into one of them. Following recent studies (Ericson et al., 2003; Paton et al., 2003; Thomas et al., 2004a, b; van Tuinen et al., 2004; Paton & Baker, 2006), the waders may be more accurately subdivided as follows:

Dunlin (Calidris alpina).
  • Suborder Scolopaci
    • Family Scolopacidae: snipe, sandpipers, phalaropes, and allies
  • Suborder Thinocori
    • Family Rostratulidae: painted snipe
    • Family Jacanidae: jacanas
    • Family Thinocoridae: seedsnipe
    • Family Pedionomidae: Plains Wanderer
  • Suborder Chionidi
    • Family Burhinidae: thick-knees
    • Family Chionididae: sheathbills
    • Family Pluvianellidae: Magellanic Plover
  • Suborder Charadrii
    • Family Ibidorhynchidae: Ibisbill
    • Family Recurvirostridae: avocets and stilts
    • Family Haematopodidae: oystercatchers
    • Family Charadriidae: plovers and lapwings

In keeping more in line with the traditional grouping, the Thinocori could be included in the Scolopaci, and the Chionidi in the Charadrii. However, the increasing knowledge about the early evolutionary history of modern birds suggests that the assumption of Paton et al. (2003) and Thomas et al. (2004b) of 4 distinct "wader" lineages (= suborders) already being present around the K-T boundary is correct.

See also


  • Paton, Tara A. & Baker, Allan J. (2006): Sequences from 14 mitochondrial genes provide a well-supported phylogeny of the Charadriiform birds congruent with the nuclear RAG-1 tree. Molecular Phylogenetics and Evolution 39(3): 657–667. doi:10.1016/j.ympev.2006.01.011 (HTML abstract)
  • Paton, T. A.; Baker, A. J.; Groth, J. G. & Barrowclough, G. F. (2003): RAG-1 sequences resolve phylogenetic relationships within charadriiform birds. Molecular Phylogenetics and Evolution 29: 268-278. doi:10.1016/S1055-7903(03)00098-8 (HTML abstract)
  • Thomas, Gavin H.; Wills, Matthew A. & Székely, Tamás (2004a): Phylogeny of shorebirds, gulls, and alcids (Aves: Charadrii) from the cytochrome-b gene: parsimony, Bayesian inference, minimum evolution, and quartet puzzling. Molecular Phylogenetics and Evolution 30(3): 516-526. doi:10.1016/S1055-7903(03)00222-7 (HTML abstract)
  • van Tuinen, Marcel; Waterhouse, David & Dyke, Gareth J. (2004): Avian molecular systematics on the rebound: a fresh look at modern shorebird phylogenetic relationships. Journal of Avian Biology 35(3): 191-194. PDF fulltext

Wikimedia Foundation. 2010.

См. также в других словарях:

  • Wader — Wader,   Hannes, Liedermacher, * Bielefeld 23. 6. 1942; wurde nach 1963 mit eigenen Liedern auf den Chanson Folklore Festivals auf der Burg Waldeck bekannt. Seit etwa 1975 interpretiert er traditionelle Volkslieder, auch auf Plattdeutsch, sowie… …   Universal-Lexikon

  • Wader — Wad er, n. 1. One who, or that which, wades. [1913 Webster] 2. (Zo[ o]l.) Any long legged bird that wades in the water in search of food, especially any species of limicoline or grallatorial birds; called also {wading bird}. See Illust. g, under… …   The Collaborative International Dictionary of English

  • Wader — Wader, in Graubünden s.v.w. Gletscher …   Kleines Konversations-Lexikon

  • wader — ► NOUN 1) a sandpiper, plover, or other wading bird. 2) (waders) high waterproof boots, used by anglers …   English terms dictionary

  • wader — [wā′dər] n. 1. a person or thing that wades 2. any bird that wades in marshes, lakes, etc. 3. a) [pl.] high waterproof boots ☆ b) [usually pl.] waterproof overalls with bootlike parts for the feet, worn as by fishermen for wading in deep water …   English World dictionary

  • Wader — Hannes Wader, 2004 Hannes Wader (* 23. Juni 1942 in Hoberge Uerentrup/Bielefeld, Nordrhein Westfalen als Hans Eckard Wader) ist ein Musiker und Liedermacher. Er gilt neben Reinhard Mey, Konstantin Wecker …   Deutsch Wikipedia

  • wader — UK [ˈweɪdə(r)] / US [ˈweɪdər] noun Word forms wader : singular wader plural waders 1) waders [plural] rubber boots or a combination of trousers and rubber boots that people wear when they stand in water to fish 2) [countable] a wading bird …   English dictionary

  • wader — [[t]we͟ɪdə(r)[/t]] waders 1) N COUNT A wader is a bird with long legs and a long neck, which lives near water and feeds on fish. There are several different kinds of waders. 2) N COUNT: usu pl Waders are long rubber boots which cover all of the… …   English dictionary

  • wader — noun Date: 1673 1. one that wades 2. shorebird; also wading bird 3. plural high waterproof boots or a one piece waterproof garment usually consisting of pants with attached boots that are used for wading (as when fishing) …   New Collegiate Dictionary

  • wader — /way deuhr/, n. 1. a person or thing that wades. 2. Also called wading bird. any of various large birds having long legs, long necks, and long bills, that are adapted for wading in shallow waters and living on fish, frogs, etc., as the crane,… …   Universalium

Поделиться ссылкой на выделенное

Прямая ссылка:
Нажмите правой клавишей мыши и выберите «Копировать ссылку»