Carpinus caroliniana

Carpinus caroliniana
American Hornbeam foliage
Scientific classification
Kingdom:	Plantae
(unranked):	Angiosperms
(unranked):	Eudicots
(unranked):	Rosids
Order:	Fagales
Family:	Betulaceae
Genus:	Carpinus
Species:	C. caroliniana
Binomial name
Carpinus caroliniana Walter

Carpinus caroliniana (American hornbeam) is a small hardwood tree in the genus Carpinus. American hornbeam is also occasionally known as blue-beech, ironwood, or musclewood. It is native to eastern North America, from Minnesota and southern Ontario east to Maine, and south to eastern Texas and northern Florida. It also grows in Canada (southwest Quebec and southeast Ontario), Mexico (central and southern), Guatemala, and western Honduras.^[1]

The bark of the American Hornbeam

It is a small tree reaching heights of 10–15 m, rarely 20 m, and often has a fluted and crooked trunk. The bark is smooth and greenish-grey, becoming shallowly fissured in old trees. The leaves are alternate, 3–12 cm long, with prominent veins giving a distinctive corrugated texture, and a serrated margin. The male and female catkins appear in spring at the same time as the leaves. The fruit is a small 7–8 mm long nut, partially surrounded by a three- to seven-pointed leafy involucre 2–3 cm long; it matures in autumn. The seeds often do not germinate till the spring of the second year after maturating.

There are two subspecies, which intergrade extensively where they meet:

• Carpinus caroliniana subsp. caroliniana. Atlantic coastal plain north to Delaware, and lower Mississippi Valley west to eastern Texas. Leaves mostly smaller, 3–9 cm long, and relatively broader, 3–6 cm broad.
• Carpinus caroliniana subsp. virginiana. Appalachian Mountains and west to Minnesota and south to Arkansas. Leaves mostly larger, 8–12 cm long, and relatively narrower, 3.5–6 cm broad.

It is a shade-loving tree, which prefers moderate soil fertility and moisture. It has a shallow, wide-spreading root system. The wood is heavy and hard, and is used for tool handles, longbows, walking sticks, walking canes and golf clubs. The leaves are eaten by the caterpillars of some Lepidoptera, for example the Io moth (Automeris io).

Description

Common along the borders of streams and swamps, loves a deep moist soil. Varies from shrub to small tree, and ranges throughout the United States east of the Rocky Mountains.

• Bark: On old trees near the base, furrowed. Young trees and branches smooth, dark bluish gray, sometimes furrowed, light and dark gray. Branchlets at first pale green, changing to reddish brown, ultimately dull gray.
• Wood: Light brown, sapwood nearly white; heavy, hard, close-grained, very strong. Used for levers, handles of tools. Sp. gr., 0.7286; weight 45.41 lbs.
• Winter buds: Ovate, acute, chestnut brown, one-eighth of an inch long. Inner scales enlarge when spring growth begins. No terminal bud is formed.
• Leaves: Alternate, two to four inches long, ovate-oblong, rounded, wedge-shaped, or rarely subcordate and often unequal at base, sharply and doubly serrate, acute or acuminate. They come out of the bud pale bronze green and hairy; when full grown they are dull deep green above, paler beneath; feather-veined, midrib and veins very prominent on under side. In autumn bright red, deep scarlet and orange. Petioles short, slender, hairy. Stipules caducous.
• Flowers: April. Monœcious, apetalous, the staminate naked in pendulous aments. The staminate ament buds are axillary and form in the autumn and during the winter resemble leaf-buds, only twice as large; these aments begin to lengthen very early in the spring, when full grown are about one and one-half inches long. The staminate flower is composed of three to twenty stamens crowded on a hairy torus, adnate to the base of a broadly ovate, acute boot-shaped scale, green below the middle, bright red at apex. The pistillate aments are one-half to three-fourths of an inch long with ovate, acute, hairy, green scales and bright scarlet styles.
• Fruit: Clusters of involucres, hanging from the ends of leafy branches. Each involucre slightly incloses a small oval nut. The involucres are short stalked, usually three-lobed, though one lobe is often wanting; halberd-shaped, coarsely serrate on one margin, or entire.^[2]

References

1. ^ "Silvics of North America, Vol.2 (USDA Forest Service)". http://na.fs.fed.us/pubs/silvics_manual/volume_2/carpinus/caroliniana.htm. 
2. ^ Keeler, Harriet L. (1900). Our Native Trees and How to Identify Them. New York: Charles Scriber's Sons. pp. 319–322.