Aspen parkland

Aspen parkland refers to a transitional biome between prairie and boreal forest stretching from northeastern British Columbia through central and northwestern Alberta, central Saskatchewan to central and southern Manitoba. Aspen parkland consists of groves of aspen poplars and spruce interspersed with areas of prairie grasslands. This ecoregion is also intersected by large stream and river valleys lined with aspen-spruce forests and dense shrubbery. This is a zone of constant competition and tension. Within the parklands, prairie and woodlands struggle to overtake each other. [citebook|title=Aboriginal People and Colonizers of Western Canada to 1900 |author= Sarah Carter |year=1999|publisher=University of Toronto Press| pages=19|id=ISBN 0802079954 |url=http://books.google.com/books?id=MhjguytskOkC&pg=PA19&ots=vEMJQVPN4V&dq=%22Aspen+parkland%22&as_brr=3&ei=Ot2-RonLIJSS7QKvxeDyCw&ie=ISO-8859-1&output=html&sig=zAV1X1TaGRC5no2LTNloxL2a3fY]

This article focuses on this biome in North America. Similar biomes also exist in Russia north of the steppes and in northern Europe.

etting

The aspen parkland biome runs in a thin band no wider than 500 km through the Prairie Provinces, although it gets broader to the west, especially in Alberta. The cities of Edmonton and Saskatoon are the largest cities completely in this biome. Winnipeg is bordered by tallgrass prairie to the west and south. Calgary is bordered by prairie to the east and the Foothills Parkland to the west.

In Alberta, there are three main sections of aspen parkland: Central, Peace River and Foothills. These sections are distinguished by their geographical location. The Central Parkland is the largest section and is part of main band of aspen parkland which extends into Saskatchewan and Manitoba. It is bordered by prairie to the south and the boreal forest to the north. The Peace River Parkland is located in the Peace River region of the province and is completely surrounded by boreal forest, cut off from the Central Parkland. It also extends into northeastern British Columbia.

Climate

The climate is continental with long cold winters and short summers which tend to be cool to warm. Seasonal temperature extremes are common, and are greatest in Saskatchewan and Manitoba. Alberta's aspen parkland is fairly moderate in seasonal extremes by comparison, however, winters can still be long and harsh. Precipitation is generally lower than in the boreal forest, but is still higher than in the prairies. Hence, groves of forests and lakes are able to thrive where they cannot do so in the prairies.

The Peace River Parkland is generally cooler than the Central Parkland, but still supports extensive farmland. The Foothills Parkland is located to the west of the grasslands in the southern section of the province and just east of the Canadian Rockies. The Foothills Parkland can get very windy. Chinooks are also common there during the winter.

Flora

Four significantly different habitats are common in the aspen parklands: The fescue prairie, the woodlands, the ravines and the wetlands and lakes. A rarer habitat type, tallgrass aspen parkland, occurs only in the extreme southeastern corner of the aspen parklands biome (southeastern Manitoba/northwestern Minnesota). The fescue prairie is a meadowland rich in vegetation variety which forms the cover for the development of the richer soils that underlie the parklands. The close association with woodlands and wetlands makes this a choice location for many plants and a preferred range or home site for a wide diversity of wildlife. The richer soil and increased precipitation favours the natural growth of fescue grass. However, varying conditions such as moisture level and grazing pressures allow for the invasion of secondary plant species.

There are numerous grasses and sedges in the fescue prairie. Gravelly and rocky terrain is a good location for parry oat grass. Dry areas favour June, porcupine and spear grass. Wet areas are often covered with slender wheat grass and timber oat grass. Prairie rose and snowberry are common shrubs found in these grasslands.

The forested, or woodlands area is dominated by the trembling aspens and/or spruce, although other species of trees do occur. Pines, mostly jack pine and lodgepole pine will often grow in areas that have sandy soil conditions. Other native species may include box elder, tamarack and willow. The proportion of forests to grasslands has increased somewhat over the prairie in areas not affected by agriculture in the last 100 years. This increase is partly due to the reduction of prairie fires which used to destroy the new saplings on the fringes of the aspen groves. Also, it was a common practice for farmers to plant stands of trees as windbreaks.

Aspen woodlands support an extensive understory consisting of mid-sized and small shrubs, some herbs and ground cover. Spruce-dominated woodlands usually do not support a dense understory due to more acidic and nutrient-poor soils and a denser canopy, which reduces sunlight reaching the forest floor below. However, in areas where a mixture of aspen and spruce occur, a fairly dense understory can still thrive. In fact, the mixed wood understory, as it is called, supports the greatest diversity of forest wildlife in the aspen parkland.

Large shrubs such as red-osier dogwood, beaked willow, saskatoon, chokecherry and pincherry, along with the smaller shrubs including prickly rose, snowberry, beaked hazelnut and low bush cranberry, form a dense entangled understory. Dense shrubbery is a typical feature in aspen-dominated forests. Common herbs found in the woodlands include: Lindley’s aster "(Aster ciliolatus)", northern bedstraw "(Galium boreale)", pea vine, Western Canada violet "(Viola canadensis)", dewberry and bunchberry. Mosses appear at the base of trees and on the ground.

Wetlands are very common in this biome, including lakes, shallow open water, marshes, and grassy wetlands. Glacial erosion has contributed to such features by creating depressions in which standing water can collect. In the larger depressions, permanent lakes or ponds of water remain. Many of the lakes have a saline character, thus most shore vegetation has a high tolerance of salty soils. These lakes are known as alkali lakes. Wet meadows are flooded in the spring and dry by fall. They contain rushes, sedges and grasses and provide excellent opportunities to study the similarities and differences of these forms of vegetation.

Rivers and streams erode valleys throughout the parkland ecoregion. Steep hills and ravines result in a unique topography. Southwest slopes with increased exposure to the sun are dry and often more grass covered while the shaded north and east exposures retain more moisture and tend to have greater forest cover. Some forms of vegetation unique to the ravines include: poplar, spruce, birch, willow, and river alder.

Wildflowers are an important component of the grassland association of the parkland. Look for common yarrow, cut-leaf anemone, rock cress, creeping white prairie aster, milk vetch, late yellow loco weed, goldenrod, prairie rose, prairie crocus, and tiger lily.

The aspen understory

There are three main factors which influence the understory vegetation in the aspen stands of the mixed wood forest.

1. Good sun exposure encourages a dense vegetation growth below the canopy. This is of particular importance in the early spring before the trees are in leaf.

2. Warm soil and air temperature at the base level result in rapid melting process in spring which favours the growth of shrubs.

3. A large percentage of precipitation passes through the canopy. This provides a protective snow cover in winter and in warm seasons precipitation percolates through the leaf cover to nourish plants which require surface soil moisture.

The result of the above factors is an extensive understory of vegetation in the aspen forest. Common shrubs and herbs are: saskatoon, red-osier dogwood, raspberry, wild rose, currants and bracted honeysuckle, wild sarsaparilla, hairy lungwort, asters, and peavine. Twinflower, strawberries, bunchberries, horsetails and wintergreen form an attractive grown cover.

The mineral soil is covered by a decaying cover of organic matter. Numerous consumers and decomposers create humus materials. Burrowing animals mix the new fertile materials with the soil to form a rich rooting compound.

The spruce forest understory

Factors which influence the understory vegetation of spruce stands in the boreal forest association include:

1. Year round reduced sun exposure below the canopy restricts the forest undergrowth to shade tolerant species.

2. A large percentage of the precipitation is trapped in the upper tree boughs of the spruce forest and is released through evaporation. The ground cover of feather moss quickly absorbs most of the moisture which does penetrate the canopy. These factors combine to cause drier conditions in the underlying mineral soils.

3. The fallen acidic spruce needles are not fully decomposed and combine with the moss base. Water held in the moss carries the acid from the spruce needles into the mineral soil and leaches out soil nutrients – leaving a highly acidic, low nutrient soil base which is unsuitable for most boreal vegetation.

As a result of the above factors the forest floor ranges from nearly devoid of vegetation to a dense carpet of feather moss. A sparse community of shade tolerant shrubs exists in this environment. Some species of plants in the understory are Green Alder, low bush cranberry, prickly rose, bunchberry, twinflower, wild lily-of-the-valley, northern Comandra and wintergreens.

The mixedwood stands understory

There are several factors which influence the mixedwood stands in the boreal forest.

Where there are stands of aspen and spruce forests in close association with each other, a mixed wood forest occurs. Each group forms its own microassociation as described previously.

When the spruce and aspen forest types are mixed, the result can be quite different from the aspen or spruce stands. Animal and vegetation associations from each type combine to create considerable diversity of habitat which is typical of either spruce or aspen stands. The mixture of the transition soils provides an attractive environment with either pure spruce or pure aspen woodlands. An example of a bird which prefers a mixed wood habitat is the yellow-rumped warbler.

The mixedwood forest wetlands consist mainly of bogs, fens and marshes. Black spruce, tamarack, willow and bog and sphagnum mosses are the major vegetation types found in these lowlands. Dwarf birch and sedges cover large, wet areas with jack pine occurring on the sandy ridges.

Bear, moose, foxes, coyotes, beaver, snowshoe hare and red squirrels are found most often in the mixedwood stands compared to the aspen forests and spruce forests. Several species of warblers find this a preferred habitat. Solitary sandpipers and Canada geese are residents of the wetlands.

Fauna

Wildlife in the parklands include moose, white-tailed deer, black bears, coyotes, pocket gophers, thirteen-lined ground squirrels, Richardson’s ground squirrels, beavers, snowshoe hares, weasels, gray wolves, kingfishers, ruffed grouse, magpies and northern orioles.

Burrowing rodents such as Richardson’s ground squirrels, thirteen-lined ground squirrels, and pocket gophers play a major role in the balance between the aspen groves and the grassland. These excavators make mounds of fresh soil which are ideal locations for the germination of poplar seeds. Once established, these trees spread by suckering, thus creating a new aspen groves.

White-tailed deer finds shelter in the aspen and graze on the grasslands; coyotes and foxes hunt the resident rodents. Historically, bison grazed on the grassland and helped to prevent the spread of aspen groves. However, bison are now mostly absent due to overhunting during settlement in the 19th Century and extensive loss of habitat due to agriculture. Bison, however, can still be seen in protected areas such as Elk Island National Park east of Edmonton and in farms, where they are raised for meat.

Extensive cultivation has disturbed the habitats of some birds which nest and feed on the fescue grassland. However, the horned lark and meadowlark have managed to adapt to the new conditions. Song sparrow, vesper sparrow, and American goldfinch can often be seen in open areas.

Wildlife in the woodlands is varied and abundant. The varying hare, weasel, fox, coyote, and white-tailed deer make their homes in this region.

Equally abundant are a variety of bird species. Black-capped chickadee, hairy woodpeckers, ruffed grouse, magpies, and great horned owls can be observed in all seasons. Summer residents include: red-eyed vireo, least flycatcher and northern oriole.

The invertebrate population in the woodland is enormous. Some of the most common invertebrates are roundworms, snails, segmented worms, centipedes, mites, spiders and mosquitoes. Poplar gore beetles and forest tent caterpillars are destructive to the tree cover.

Water dependent mammals who make the ravines and wetlands their home are beaver, muskrats, otters and mink. Birds which prefer the wetland habitat include kingfishers and bank swallows. Insects in this region include caddis flies, mayflies and black flies.

There is an abundance of bird life around the wetland marshes. Many species of ducks make their summer homes in these waters and Canada geese nest in the more remote marshes. Blackbirds, marsh wrens and black terns nest in the reeds. Franklin gulls nest in the marsh vegetation, but range over agricultural fields for grasshoppers, crickets, and mice. Shore birds include: avocet, piping plover, spotted sandpiper, willet, common snipe and killdeer.

Human use and conservation

Before European colonization, there were large areas of western aspen and aspen parkland in the west of what would become Canada and the United States. This was maintained by light to moderate fires with a frequency of 3 to 15 years. Fire also swept the Rocky Mountains aspen as frequently as every ten years, creating large areas of parkland. Settlement increased fire frequency in the late 1800s until fire suppression became popular.cite web|url=http://www.treesearch.fs.fed.us/pubs/4554|title=Wildland fire in ecosystems: effects of fire on flora|last=Brown|first=James K.|coauthors=Smith, Jane Kapler|date=2000|work=Gen. Tech. Rep. RMRS-GTR-42-vol. 2|publisher=Department of Agriculture, Forest Service, Rocky Mountain Research Station|pages=40|accessdate=2008-07-20]

Most of the aspen parkland, like the prairie biome, has been extensively altered by agriculture over the last 100 years since settlement first began in the late 19th Century. While the climate is generally cooler than in the prairies, the climate is still mild and dry enough to support large-scale farming. The soils in the aspen parkland biome are also quite fertile, especially around Edmonton and Saskatoon. Oil and natural gas exploration and drilling have also disturbed the natural habitat as well, especially in Alberta and northeastern British Columbia. As a result, undisturbed habitat is now no longer as nearly common as it was a century ago.

References

External links

* [http://www.worldwildlife.org/wildworld/profiles/terrestrial/na/na0802_full.html Canadian Aspen forests and parklands (World Wildlife Fund)]
* [http://www.cas.vanderbilt.edu/bioimages/ecoregions/50802.htm Canadian Aspen forests and parklands (Vanderbilt University)]