Biogeochemical cycle

A commonly cited example is the water cycle.

In ecology and Earth science, a biogeochemical cycle or substance turnover or cycling of substances is a pathway by which a chemical element or molecule moves through both biotic (biosphere) and abiotic (lithosphere, atmosphere, and hydrosphere) compartments of Earth. A cycle is a series of change which comes back to the starting point and which can be repeated. The term “biogeochemical” tells us that biological; geological and chemical factors are all involved. On the other hand the circulation of chemical nutrients like carbon, oxygen, nitrogen, phosphorus, calcium, and water etc. through the biological and physical world are known as biogeochemical cycle. In effect, the element is recycled, although in some cycles there may be places (called reservoirs) where the element is accumulated or held for a long period of time (such as an ocean or lake for water). Water, for example, is always recycled through the water cycle, as shown in the diagram. The water undergoes evaporation, condensation, and precipitation, falling back to Earth clean and fresh. Elements, chemical compounds, and other forms of matter are passed from one organism to another and from one part of the biosphere to another through the biogeochemical cycles.^[1][2]

Systems

Chloroplasts conduct photosynthesis and are found in plant cells and other eukaryotic organisms. These are Chloroplasts visible in the cells of Plagiomnium affine — Many-fruited Thyme-moss.

Ecological systems (ecosystems) have many biogeochemical cycles operating as a part of the system, for example the water cycle, the carbon cycle, the nitrogen cycle, etc. All chemical elements occurring in organisms are part of biogeochemical cycles. In addition to being a part of living organisms, these chemical elements also cycle through abiotic factors of ecosystems such as water (hydrosphere), land (lithosphere), and/or the air (atmosphere). The living factors of the planet can be referred to collectively as the biosphere. All the nutrients—such as carbon, nitrogen, oxygen, phosphorus, and sulfur—used in ecosystems by living organisms are a part of a closed system; therefore, these chemicals are recycled instead of being lost and replenished constantly such as in an open system.^[3]

The flow of energy in an ecosystem is an open system; the sun constantly gives the planet energy in the form of light while it is eventually used and lost in the form of heat throughout the trophic levels of a food web. Carbon is used to make carbohydrates, fats, and proteins, the major sources of food energy. These compounds are oxidized to release carbon dioxide, which can be captured by plants to make organic compounds. The chemical reaction is powered by the light energy of the sun.

It is possible for an ecosystem to obtain energy without sunlight. Carbon must be combined with hydrogen and oxygen in order to be utilized as an energy source, and this process depends on sunlight. Ecosystems in the deep sea, where no sunlight can penetrate, use sulfur. Hydrogen sulfide near hydrothermal vents can be utilized by organisms such as the giant tube worm. In the sulfur cycle, sulfur can be forever recycled as a source of energy. Energy can be released through the oxidation and reduction of sulfur compounds (e.g., oxidizing elemental sulfur to sulfite and then to sulfate).

Although the Earth constantly receives energy from the sun, its chemical composition is essentially fixed, as additional matter is only occasionally added by meteorites. Because this chemical composition is not replenished like energy, all processes that depend on these chemicals must be recycled. These cycles include both the living biosphere and the nonliving lithosphere, atmosphere, and hydrosphere.

Reservoirs

Coal is a reservoir of carbon

The chemicals are sometimes held for long periods of time in one place. This place is called a reservoir, which, for example, includes such things as coal deposits that are storing carbon for a long period of time.^[4] When chemicals are held for only short periods of time, they are being held in exchange pools. Examples of exchange pools include plants and animals.^[4] Plants and animals utilize carbon to produce carbohydrates, fats, and proteins, which can then be used to build their internal structures or to obtain energy. Plants and animals temporarily use carbon in their systems and then release it back into the air or surrounding medium. Generally, reservoirs are abiotic factors whereas exchange pools are biotic factors.^[5] Carbon is held for a relatively short time in plants and animals in comparison to coal deposits. The amount of time that a chemical is held in one place is called its residence.^[4]

Important cycles

The most well-known and important biogeochemical cycles, for example, include the carbon cycle, the nitrogen cycle, the oxygen cycle, the phosphorus cycle, the sulfur cycle and the water cycle. There are many biogeochemical cycles that are currently being studied for the first time as climate change and human impacts are drastically changing the speed, intensity, and balance of these relatively unknown cycles. These newly studied biogeochemical cycles include the mercury cycle^[6] and the human-caused cycle of atrazine, which may affect certain species.

Biogeochemical cycles always involve hot equilibrium states: a balance in the cycling of the element between compartments. However, overall balance may involve compartments distributed on a global scale.

As biogeochemical cycles describe the movements of substances on the entire globe, the study of these is inherently multidiciplinary. The carbon cycle may be related to research in ecology and atmospheric sciences.^[7] Biochemical dynamics would also be related to the fields of geology and pedology (soil study).^[8]

Global biogeochemical cycles critical for life

Nitrogen cycle   Water cycle   Carbon cycle   Oxygen cycle   Phosphorus cycle

See also

	Environment portal
	Ecology portal
	Earth sciences portal
	Sustainable development portal

• Biodiversity
• Bioregion
• Earth Science
• Ecology
• Ecosystem
• Natural environment
• Nature
• Recycling (ecological)
• Sulfur Cycle

References

Modelling ecosystems – trophic components General Abiotic component Abiotic stress Behaviour Biogeochemical cycle Biomass Biotic component Biotic stress Carrying capacity Competition Ecosystem Ecosystem ecology Ecosystem model Keystone species List of feeding behaviours Metabolic theory of ecology Productivity Producers Autotrophs Chemosynthesis Chemotrophs Foundation species Mixotrophs Myco-heterotrophy Mycotroph Organotrophs Photoheterotrophs Photosynthesis Photosynthetic efficiency Phototrophs Primary nutritional groups Primary production Consumers Apex predator Bacterivore Carnivores Chemoorganotroph Foraging Generalist and specialist species Intraguild predation Herbivores Heterotroph Heterotrophic nutrition Insectivore Mesopredator release hypothesis Omnivores Optimal foraging theory Predation Prey switching Decomposers Chemoorganoheterotrophy Decomposition Detritivores Detritus Microorganisms Archaea Bacteriophage Environmental microbiology Lithoautotroph Lithotrophy Microbial cooperation Microbial ecology Microbial food web Microbial intelligence Microbial loop Microbial mat Microbial metabolism Phage ecology Food webs Biomagnification Ecological efficiency Ecological pyramid Energy flow Food chain Trophic level Example webs Cold seeps Hydrothermal vents Intertidal Kelp forests Lakes North Pacific Subtropical Gyre Rivers San Francisco Estuary Soil Tidal pool Processes Ascendency Bioaccumulation Cascade effect Climax community Competitive exclusion principle Consumer-resource systems Copiotrophs Dominance Ecological network Ecological succession Energy quality Energy Systems Language f-ratio Feed conversion ratio Feeding frenzy Mesotrophic soil Nutrient cycle Oligotroph Paradox of the plankton Trophic cascade Trophic mutualism Trophic state index Defense/counter Animal coloration Antipredator adaptations Herbivore adaptations to plant defense Mimicry Plant defense against herbivory Predator avoidance in schooling fish  Modelling ecosystems – other components Population ecology Abundance Allee effect Depensation Ecological yield Effective population size Intraspecific competition Logistic function Malthusian growth model Maximum sustainable yield Overpopulation in wild animals Overexploitation Population cycle Population dynamics Population modeling Population size Predator–prey equations Recruitment Resilience Small population size Stability Species Biodiversity Density-dependent inhibition Ecological effects of biodiversity Ecological extinction Endemic species Flagship species Gradient analysis Indicator species Introduced species Invasive species Latitudinal gradients in species diversity Minimum viable population Neutral theory Occupancy-abundance relationship Population viability analysis Priority effect Rapoport's rule Relative abundance distribution Relative species abundance Species diversity Species homogeneity Species richness Species distribution Species-area curve Umbrella species Species interaction Antibiosis Biological interaction Commensalism Community ecology Ecological facilitation Interspecific competition Mutualism Storage effect Spatial ecology Biogeography Cross-boundary subsidy Ecocline Ecotone Ecotype Disturbance Edge effect Foster's rule Habitat fragmentation Ideal free distribution Intermediate Disturbance Hypothesis Island biogeography Landscape ecology Landscape epidemiology Landscape limnology Metapopulation Patch dynamics r/K selection theory Source–sink dynamics Niche Ecological niche Ecological trap Ecosystem engineer Environmental niche modelling Guild Habitat Limiting similarity Niche apportionment models Niche construction Niche differentiation Other networks Assembly rules Bateman's principle Bioluminescence Ecological collapse Ecological debt Ecological deficit Ecological energetics Ecological indicator Ecological threshold Ecosystem diversity Emergence Extinction debt Kleiber's law Liebig's law of the minimum Marginal value theorem Thorson's rule Xerosere Other Allometry Alternative stable state Balance of nature Biological data visualization Constructal theory Ecocline Ecological economics Ecological footprint Ecological forecasting Ecological humanities Ecological stoichiometry Ecopath Ecosystem based fisheries Endolith Evolutionary ecology Functional ecology Industrial ecology Macroecology Microecosystem Natural environment Systems ecology Theoretical ecology List of ecology topics