Pharyngeal arch

Infobox Embryology
Name = Branchial arch
Latin =
GraySubject = 13
GrayPage = 65


Caption = Schematic of developing fetus with first, second and third arches labeled.


Caption2 = Floor of pharynx of human embryo about twenty-six days old.
System =
CarnegieStage = 10
Precursor =
GivesRiseTo =
MeshName = Branchial+Arches
MeshNumber = A16.254.160
DorlandsPre = a_57
DorlandsSuf = 12149648
In the development of vertebrate animals, the pharyngeal arches (also called branchial arches or gill arches in fish) develop during the fourth and fifth week "in utero" as a series of mesodermal outpouchings on the left and right sides of the developing pharynx. In fish, the branchial arches give rise to gills.

Development

These grow and join in the ventral midline. The first arch, as the first to form, separates the mouth pit or stomodeum from the pericardium. By differential growth the neck elongates and new arches form, so the pharynx has six arches ultimately.

Each pharyngeal arch has a cartilaginous bar, a muscle component which differentiates from the cartilagenous tissue, an artery, and a cranial nerve.Each of these is surrounded by Mesenchyme. Arches do not develop simultaneously, but instead possess a "staggered" development.

Relations

Pharyngeal or branchial pouches form on the endodermal side between the arches, and pharyngeal grooves (or clefts) form from the lateral ectodermal surface of the neck region to separate the arches. [cite web |url=http://www.med.howard.edu/anatomy/gas/wk7/Lecture%2024.htm |title=Lecture 24. Branchial Apparatus |accessdate=2007-09-09 |format= |work=]

The pouches line up with the clefts, and these thin segments become gills in fish.

In mammals the endoderm and ectoderm not only remain intact, but continue to be separated by a mesoderm layer.

pecific arches

There are six pharyngeal arches, but in humans the fifth arch only exists transiently during embryologic growth and development. Since no human structures result from the fifth arch, the arches in humans are I, II, III, IV, and VI. [cite web |url=http://isc.temple.edu/marino/embryology/parch98/parch_text.htm |title=Text for Pharyngeal Arch Development |accessdate=2007-09-09 |format= |work=]

More is known about the fate of the first arch than the remaining four. The first three contribute to structures above the larynx, while the last two contribute to the larynx and trachea.

Use in staging

The development of the pharyngeal arches provide a useful morphological landmark with which to establish the precise stage of embryonic development. Their formation and development corresponds to Carnegie stages 10 to 16 in mammals, and Hamburger-Hamilton stages 14 to 28 in the chicken.

ee also

* Aortic arches
* Branchial pouch
* Branchial cleft cyst

References

External links

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